Putative graviperception mechanisms of protists.

Many (if not all) free-living cells use the gravity vector for their spatial orientation (gravitaxis). Additional responses may include gravikinesis as well as changes in morphological and physiological parameters. Though using essentially different modes of locomotion, ameboid and ciliated cells seem to rely on common fundamental graviperception mechanisms. Uniquely in the ciliate family Loxodidae a specialized intracellular gravireceptor organelle has been developed, whereas in all other cells common cell structures seem to be responsible for gravisensing. Changes in direction or magnitude of acceleration (from 0 to 5 g) as well as experiments in density-adjusted media strongly indicate that either the whole cytoplasm or dense organelles like nuclei act as statoliths and open directly or via cytoskeletal elements mechano-sensitive ion channels in the cell membrane. A recent spaceflight experiment (S/MM-06) demonstrated that prolonged (9 d) actual weightlessness did not affect the ability of Loxodes to respond to acceleration stimuli. However, prolonged cooling (> or = l4 d, 4-10 degrees C) destroyed the ability for gravitactic orientation of Paramecium. This may reflect a profound effect either on the gravireceptor itself or on the gravity-signal processing. In gravity signalling the ubiquitous second messenger cAMP may be involved in acceleration-stimulus transduction.     

Microperturbations on the <Emphasis Type="Italic">International Space Station</Emphasis> during physical exercises of the crew

The results of a preliminary analysis of microperturbations on the International Space Station during physical exercises of the crew are presented. The goal of this paper is to identify the parameters of perturbations when physical exercises are performed. The results of measurements by sensors of microaccelerations of both Russian and American segments during physical exercises in the service module of the Russian segment are analyzed.     

Optical properties of the undisturbed mesosphere from wide-angle twilight sky polarimetry

The paper describes the first results of all-sky polarization measurements of the twilight background started in central Russia in the very beginning of summer 2011. Time-frequency data of the sky intensity and polarization over a wide range of sky point zenith distances are used to separate single and multiple scattering and construct the altitude dependence of the scattering coefficient and polarization in the mesosphere (altitudes from 60 to 90 km) at different angles. The undisturbed structure of the mesosphere without noticeable aerosol stratification on observation days makes it possible to estimate the temperature of the atmosphere at these altitudes.     

A method to ensure energy security of satellite communication systems

A method is substantiated to ensure energy security for the satellite communication systems (SCS) at a close position of the radio interception receiver. This is done by lowering the carrier frequency down to f ₀ = 60…80 MHz and by applying spaced measurements with n ≥ 4 receiving antennas.     

Stellar coronagraph using the principle of achromatic null-interferometer

In order to observe exoplanets we propose a space-based achromatic stellar coronagraph combined with a 0.8–1.5 m telescope. We develop an achromatic common path interferometer for observing an exoplanet (a faint off-axis source) on the background of a hoste star (bright axial source). An image of the star and its copy acquire an achromatic phase shift by 180° and interfere in antiphase. The achromatic phase shift is caused by geometric phase in the scheme of a three-dimensional interferometer. The interference process divides spatially the dark and light fields of the star image redirecting them to the opposite sides of a beam splitter. The interference process does not weaken the image of a planet, with equal intensities it is redirected to both sides of a beam splitter. The suggested scheme of common path interferometer ensures mechanical stability. The background signal is experimentally demonstrated to be reduced by six orders of magnitude.     

Evolution of salt finger convection with steady wind shear

In oceanographics situations where salt fingers may be an important mechanism for the transport of heat and salt in the vertical direction, velocity shears may also be present. Salt finger convection is analogous to Bénard convection in that the kinetic energy of the motions is obtained from the potential energy stored in the unstable distribution of a stratifying component. On the basis of the thermal analogy it is of interest to discover whether salt fingers are converted into two-dimensional sheets by wind shear, and how the vertical fluxes of heat and salt are changed by wind shear. Salt finger convection under the effect of steady wind shear is theoretically examined in this paper. The evolution of instability developing in the presence of a vertical density gradient disturbance and the horizontal Couette flow is considered near the onset of salt fingers under a moderate rate of shear. We use velocity as the basic variable and solve the pressure Poisson equation in terms of the associated Green function. Growth competition between the longitudinal rolls (LR) and the transverse rolls (TR), whose axes are, respectively, in the direction parallel to and perpendicular to the Couette flow, is investigated by the weakly nonlinear analysis of coupled-mode equations. The results show that the TR mode is stable under a small wind shear and the LR mode is stable for a higher wind shear.     

Solution to the problem of stability of regular precessions of a symmetrical gyrostat in Newtonian field

The constraints imposed on parameters of a force field and of a symmetrical gyrostat in it are determined, at which regular precession of this gyrostat demonstrate Lyapunov stability (or, respectively, instability).     

Gravitropism of basidiomycetous fungi--on Earth and in microgravity.

In order to achieve perfect positioning of their lamellae for spore dispersal, fruiting bodies of higher fungi rely on the omnipresent force gravity. Only accurate negatively gravitropic orientation of the fruiting body cap will guarantee successful reproduction. A spaceflight experiment during the STS-55 Spacelab mission in 1993 confirmed that the factor gravity is employed for spatial orientation. Most likely every hypha in the transition zone between the stipe and the cap region is capable of sensing gravity. Sensing presumably involves slight sedimentation of nuclei which subsequently causes deformation of the net-like arrangement of F-actin filament strands. Hyphal elongation is probably driven by hormone-controlled activation and redistribution of vesicle traffic and vesicle incorporation into the vacuoles and cell walls to subsequently cause increased water uptake and turgor pressure. Stipe bending is achieved by way of differential growth of the flanks of the upper-most stipe region. After reorientation to a horizontal position, elongation of the upper flank hyphae decreases 40% while elongation of the lower flank slightly increases. On the cellular level gravity-stimulated vesicle accumulation was observed in hyphae of the lower flank.     

Consequences of collisions of natural cosmic bodies with the Earth’s atmosphere and surface

Possible consequences of collisions of natural cosmic bodies with the Earth’s atmosphere and surface are described. The methodological basis of classification of consequences is the solution of meteor physics equations characterizing the trajectory of a body in the atmosphere, namely, the dependence of the body’s velocity and mass on the flight altitude. The solution depends on two dimensionless parameters characterizing the drag altitude and the role of mass loss by a meteoroid during its motion in the atmosphere. Depending on values of these parameters, the degree of effect on the planetary surface considerably changes. In particular, the conditions of cratering and meteorite fall on the planetary surface are obtained. The results are presented in a simple analytical form. They quite match to the real events considered in the paper. Recommendations are given on further investigations into the important problem of interaction of cosmic bodies with planetary atmospheres.     

Wave resistance defect in a planar flow over periodic reliefs

The dependence of the wave resistance coefficients for planar periodic reliefs on the similarity parameters is investigated. It is proved that the wave resistance coefficients of the infinite reliefs and their finite analogs in the case of the whole wave numbers coincide, whereas in the case of the fractional wave numbers they differ.