Protozoa as model systems for the study of cellular responses to altered gravity conditions.

The orientation behavior of Paramecium changed in a similar way after transition to conditions of free-fall in a sounding rocket and after transition to conditions of simulated weightlessness on a fast rotating clinostat. After a period of residual orientation, Paramecium cells distributed themselves randomly 80 s (120 s) after onset of free-fall (simulated weightlessness). Swimming velocity increased significantly; however, the increase was transient and subsided after 3 min in the rocket experiments, while the velocity remained enhanced even during 2 h of rotation on a fast clinostat. Trichocysts were present and without morphological changes in Paramecium cells which had been exposed to a rocket flight, as well as to fast or slow rotation on a clinostat. Regeneration of the oral apparatus of Stentor and morphogenesis of Eufolliculina proceeded normally on the clinostat. The results demonstrate that the clinostat is a useful tool to simulate the conditions of weightlessness on earth and to detect gravisensitive cellular functions.     

The Origin of Solar Energetic Particle Events: Coronal Acceleration versus Shock Wave Acceleration

We review evidence that led to the view that acceleration at shock waves driven by coronal mass ejections (CMEs) is responsible for large particle events detected at 1 AU. It appears that even if the CME bow shock acceleration is a possible model for the origin of rather low energy ions, it faces difficulties on account of the production of ions far above 1 MeV: (i) although shock waves have been demonstrated to accelerate ions to energies of some MeV nucl^–1 in the interplanetary medium, their ability to achieve relativistic energies in the solar environment is unproven; (ii) SEP events producing particle enhancements at energies 100 MeV are also accompanied by flares; those accompanied only by fast CMEs have no proton signatures above 50 MeV. We emphasize detailed studies of individual high energy particle events which provide strong evidence that time-extended particle acceleration which occurs in the corona after the impulsive flare contributes to particle fluxes in space. It appears thus that the CME bow shock scenario has been overvalued and that long lasting coronal energy release processes have to be taken into account when searching for the origin of high energy SEP events.